Author Contributions: M.W., E.A.B., and S.J.S. designed research; M.W., E.A.B., and C.W.K. performed research; M.W., E.A.B., and S.J.S. analyzed data and wrote the paper.
Authors report no conflict of interest.
National Institutes of Health R01 NS084844 (S.J.S), National Science Foundation Grant 1456912 (S.J.S), Woodruff Foundation (M.W.).