- Received September 22, 2016.
- Revision received February 7, 2017.
- Accepted February 9, 2017.
- Published online February 15, 2017.
Author contributions: A.C., C.S., and D.A.P. designed research; A.C., C.S., and D.A.P. performed research; A.C., C.S., and G.O. analyzed data; A.C. and D.A.P. wrote the paper.
The authors declare no competing financial interests.
This work was supported by National Institutes of Health Grants NS060996, NS070911, DA031367, and NS094390; the Mallinckrodt Foundation; the Rita Allen Foundation; and the Brain and Behavior Research Foundation.
Rita Allen Foundation100001447
Brain and Behavior Research Foundation (Brain & Behavior Research Foundation)100000874